A horse.
“They scorned the coach, they scorned the rails,
Two spanking tits with streaming tails.”
The End of All Things.
“What spurres need now for an untamed titt.”
Barnefield: Affectionate Shepherd (1594).
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Entry taken from Dictionary of Phrase and Fable, edited by the Rev. E. Cobham Brewer, LL.D. and revised in 1895.